| EDUCATION | |
| 1989 | MD and PhD (Molecular Biology and Genetics), Johns Hopkins University School of Medicine |
| 1981 | AB (Biochemical Sciences), summa cum laude, Harvard College |
| PROFESSIONAL APPOINTMENTS | |
| 2017 | Professor, The Jackson Laboratory |
| 2017 | Professor, University of Connecticut School of Medicine |
| 2017 | Professor Emeritus, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 2001 | Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 1997 | Associate Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 1991 | Assistant Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 1989 | Staff Associate, Carnegie Institution of Washington, Department of Embryology |
| 1981 | Medical Scientist Training Program, Johns Hopkins University School of Medicine |
| HONORS | |
| 2017 | Presidential Distinguished Professor, University of Connecticut School of Medicine |
| 2015 | Fellow, National Academy of Inventors |
| 2013 | Endowed Professorship: The Michael and Ann Hankin and Partners of Brown Advisory Professor in Scientific Innovation, Johns Hopkins University School of Medicine |
| 2013 | Ho-Am Prize in Medicine |
| 2013 | Rolf Luft Award (Karolinska Institutet) |
| 2012 | Member (elected), National Academy of Sciences |
| 2010 | Fellow (elected), American Association for the Advancement of Science (AAAS) |
| 1981 | summa cum laude |
| 1980 | Phi Beta Kappa |
