EDUCATION
1989 MD and PhD (Molecular Biology and Genetics), Johns Hopkins University School of Medicine
1981 AB (Biochemical Sciences), summa cum laude, Harvard College
PROFESSIONAL APPOINTMENTS
2017 Professor, The Jackson Laboratory
2017 Professor, University of Connecticut School of Medicine
2017 Professor Emeritus, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics
2001 Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics
1997 Associate Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics
1991 Assistant Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics
1989 Staff Associate, Carnegie Institution of Washington, Department of Embryology
1981 Medical Scientist Training Program, Johns Hopkins University School of Medicine
HONORS
2017 Presidential Distinguished Professor, University of Connecticut School of Medicine
2015 Fellow, National Academy of Inventors
2013 Endowed Professorship: The Michael and Ann Hankin and Partners of Brown Advisory Professor in Scientific Innovation, Johns Hopkins University School of Medicine
2013 Ho-Am Prize in Medicine
2013 Rolf Luft Award (Karolinska Institutet)
2012 Member (elected), National Academy of Sciences
2010 Fellow (elected), American Association for the Advancement of Science (AAAS)
1981 summa cum laude
1980 Phi Beta Kappa

Curriculum Vitae