| EDUCATION |
| 1989 |
MD and PhD (Molecular Biology and Genetics), Johns Hopkins University School of Medicine |
| 1981 |
AB (Biochemical Sciences), summa cum laude, Harvard College |
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| PROFESSIONAL APPOINTMENTS |
| 2017 |
Professor, The Jackson Laboratory |
| 2017 |
Professor, University of Connecticut School of Medicine |
| 2017 |
Professor Emeritus, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 2001 |
Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 1997 |
Associate Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 1991 |
Assistant Professor, Johns Hopkins University School of Medicine, Department of Molecular Biology and Genetics |
| 1989 |
Staff Associate, Carnegie Institution of Washington, Department of Embryology |
| 1981 |
Medical Scientist Training Program, Johns Hopkins University School of Medicine |
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| HONORS |
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| 2017 |
Presidential Distinguished Professor, University of Connecticut School of Medicine |
| 2015 |
Fellow, National Academy of Inventors |
| 2013 |
Endowed Professorship: The Michael and Ann Hankin and Partners of Brown Advisory Professor in Scientific Innovation, Johns Hopkins University School of Medicine |
| 2013 |
Ho-Am Prize in Medicine |
| 2013 |
Rolf Luft Award (Karolinska Institutet) |
| 2012 |
Member (elected), National Academy of Sciences |
| 2010 |
Fellow (elected), American Association for the Advancement of Science (AAAS) |
| 1981 |
summa cum laude |
| 1980 |
Phi Beta Kappa |
